We can, however, introduce an additional variable D, called the coefficient of linkage disequilibrium, and define D to be the difference between the gametic frequency pAB and what this...We can, however, introduce an additional variable D, called the coefficient of linkage disequilibrium, and define D to be the difference between the gametic frequency pAB and what this gametic frequency would be if the loci were in linkage equilibrium: pAB=pApB+D Using pAB+pAb=pA to eliminate pAB in (5.4.3), we obtain pAb=pApb−D Likewise, using pAB+paB=pB \[p_{a B}=p_{a} p_{B}-D \nonumber \…